While exploring Fontaine in Genshin Impact 4.0 update, you will come across Hydroculus that you can collect to level up Statue of the Seven in the region and earn rewards. You will come across them while completing quests, achievements, daily commissions, etc.; however, most of them will be difficult to find on your own.
Also, it will be a tiring task if you explore the complete area looking for them, and thus, we have listed all the locations of Hydroculus in Fontaine to save you time and energy.
Genshin Impact: All Hydroculus Locations In Fontaine
There are a total of 85 Hydroculus that you can find in the Fontaine, and you can find them on the ground or in the underground area. You can find all of its locations marked on the map below:
You can travel fast from one location to another by using Teleport Waypoints and the Statue of Seven to collect them quickly. In addition to this, make sure that your HP is enough to fight enemies, as you might encounter some while exploring.